Niles and Daphne flee her wedding altar but soon discover the heavy price to pay for admitting their true love as jilted groom Donny and bitter wife Mel seek revenge.
The tension of Donnie's lawsuit and Mel's demand that Niles and Daphne not be seen in public together erupts in a major fight among Frasier, Niles and Daphne.
Frasier uses his father as a pawn in order to date the attractive owner of a home for senior citizens. Niles and Daphne plan a picnic during which they watch a meteor shower, but are accidentally separated at the crucial moment.
Roz introduces Frasier to her new boyfriend, Luke, and the two men hit it off. When Roz and Luke break up, Frasier, hides his continuing friendship with him while supporting Roz during her heartbreak.
Frasier butters up the local morning TV host to let him co-host the Seattle Christmas parade with her, but when she comes down with food poisoning (from his cooking, perhaps?), he learns that Dr. Mary will be his new partner.
Frasier receives the Lifetime Achievement Award from the Seattle Broadcasting Awards. Daphne is literally gaining weight by the minute, which is obvious to everyone but herself and Niles.
Frasier and Niles realize their helplessness when Frasier's car stalls, so they enroll in an auto repair night class. They quickly fall behind, and understand what it's like to be remedial students.
Frasier and Niles try to reawaken the career of Jackson Hedley, an old actor they saw perform Hamlet when they were young. When they see him perform again, however, they realize that he is talentless and will humiliate himself.
Frasier is supposed to go to a speed-dating event and asks Roz if he should wear a suit or sweater, remarking that the slightest decision can have far-reaching effects. We see what would have happened in each circumstance.
Kenny arrives at Frasier's place in a panic after unwittingly making a date with a woman even though he's married. Niles finally notices Daphne's weight gain.
Frasier receives tickets to the Sonics game and goes with Martin and Niles. Niles is selected to attempt the halftime half-court shot, and miraculously makes it, drinking in the adoration of Martin and the rest of Seattle.
Roz gets the green light to put on a special radio show about outer space, and agrees to let Frasier narrate. However, the two fail to come to grips with their reversed roles.
Frasier and Niles throw a fancy party to try to convince Penelope, the trustee of a foundation, to donate money to the failing prep school they attended as children.
Even after Daphne's premonition of Niles getting hurt comes true, he says he needs scientific proof before he will believe in her abilities. A psychic evaluator is hired to put her skills to the test.
While shopping for a DVD player, Frasier meets an attractive woman called Claire. He soon discovers that she is a friend of Lana, former prom queen of his school whom he dated briefly in the past.
Frasier spends the day with Lana helping her sell a house. Niles spends the day with Daphne and Eddie in the dog park. And Martin spends the day dealing with an old problem.
Frasier books a romantic weekend in Belize with Claire. On the drive to the airport, Lana suggests that he doesn't know how to be happy, but he completely disagrees with her.