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- DirectorKevin McKiddStarsEllen PompeoJustin ChambersChandra WilsonThe doctors from Grey Sloan Memorial Hospital spring into action to save the lives of two girls with a powerful story. Meanwhile, Meredith tries to cope with the fact that Amelia is now living with her, and Jackson prepares for April's return back to Seattle.
- DirectorDebbie AllenStarsEllen PompeoJustin ChambersChandra WilsonThe pressure is on as a determined Bailey tackles her first day as Chief; meanwhile, April's marriage is in jeopardy and she finds herself unable to address her problems; Meredith struggles to juggle all her responsibilities and Amelia works to define her relationship with Owen.
- DirectorRob CornStarsEllen PompeoJustin ChambersChandra WilsonWhen Maggie gets an invitation to her ex-boyfriend's wedding, she starts to reevaluate her choices.
- DirectorNicole RubioStarsEllen PompeoJustin ChambersChandra WilsonOwen suggests the interns learn about breaking bad news to patients' families.
- DirectorDebbie AllenStarsEllen PompeoJustin ChambersChandra WilsonMaggie leaves her dinner party and races to the hospital, so April takes over the cooking duties in an attempt to avoid Jackson. Meanwhile, Arizona is extremely anxious about meeting Callie's new girlfriend, who happens to be a familiar face in Meredith's life.
- DirectorJeannot SzwarcStarsEllen PompeoJustin ChambersChandra WilsonPenny's first day gets off to a rocky start. April brings a young patient from overseas with tumors on his hands. Jo and Stephanie's feud continues.
- DirectorGeary McLeodStarsEllen PompeoJustin ChambersChandra WilsonA high-stakes case is tackled by a team of doctors, and the atmosphere grows tense. Meanwhile, Bailey pressures Ben to evict their roommate; and Arizona enthusiastically reenters the dating scene with a new wingman.
- DirectorRob CornStarsEllen PompeoJustin ChambersChandra WilsonAs the rumor mill around the hospital continues to spiral out of control, Grey Sloan Memorial is flooded with injured firefighters from a nearby wildfire. Maggie struggles to keep things with Andrew professional at work; meanwhile, Jo questions Alex's priorities. Also, Alex decides to make a big step in his life.
- DirectorDenzel WashingtonStarsEllen PompeoJustin ChambersChandra WilsonMeredith is attacked by a patient, and Penny finds her injured. The staff of Grey Sloan Memorial is standing by Meredith trying to save her life.
- DirectorKevin Rodney SullivanStarsEllen PompeoJustin ChambersChandra WilsonAs the hospital deals with Meredith's trauma, Callie and Maggie perform a risky, experimental surgery on Alex's 15-year-old patient against his professional opinion. Meanwhile, Owen and Nathan are still at odds with each other, and Maggie and Andrew's relationship continues to develop.
- DirectorRob CornStarsEllen PompeoJustin ChambersChandra WilsonFlashbacks reveal the evolution of April and Jackson's relationship, beginning with their initial meeting, and the years-long process of one of Jackson's patients. Also, April finds out something life changing.
- DirectorChandra WilsonStarsEllen PompeoJustin ChambersChandra WilsonA patient from Meredith's first surgery as an intern is re-admitted to the hospital with an aneurysm; Maggie thinks Richard knows about her relationship with Andrew; Arizona considers dating again.
- DirectorCharlotte BrändströmStarsEllen PompeoJustin ChambersChandra WilsonMeredith, Jo, Bailey, Callie and Jackson head to a Veterans hospital for an spectacular yet unusual surgery. Meanwhile, a group of snobby cheerleaders come into the ER and Warren makes a risky decision.
- DirectorKevin McKiddStarsEllen PompeoJustin ChambersChandra WilsonRichard swaps pairings among the residents and attendings, baffling everyone. Meanwhile, Arizona makes a bold decision as she treats a woman who's carrying quadruplets; April faces a tough choice.
- DirectorNicole RubioStarsEllen PompeoJustin ChambersChandra WilsonJackson learns about April's pregnancy, while Alex and the team of doctors handles a long triple-organ transplant surgery. Meredith considers dating again, and Andrew insists on not receiving special treatment after he and Maggie go public.
- DirectorEric LaneuvilleStarsEllen PompeoJustin ChambersChandra WilsonMeredith takes a step forward with Will but quickly regrets it; Maggie senses a disconnection with Andrew; and Amelia reevaluates her relationship with Owen. Meanwhile, Catherine comes into town and really shakes things up with April and Jackson
- DirectorChandra WilsonStarsEllen PompeoJustin ChambersChandra WilsonApril and Arizona find themselves arguing in switched positions. Richard tries to stop Catherine interfering in others' lives. Meanwhile residents are motivated to compete for a great research grant.
- DirectorJeannot SzwarcStarsEllen PompeoJustin ChambersChandra WilsonAfter a missing child forces the hospital into lockdown, Ben decides to operate on a pregnant mother without the proper surgical equipment. Miranda, as Chief, is faced with the repercussions of the event.
- 2005– 43mTV-148.1 (953)TV EpisodeDirectorChris HaydenStarsEllen PompeoJustin ChambersChandra WilsonMiranda is faced with the repercussions of Ben's actions; Miranda forms an advisory committee
- DirectorZetna FuentesStarsEllen PompeoJustin ChambersChandra WilsonThe doctors work frantically to save a young boy who was accidentally shot. Meanwhile, Arizona is upset when Callie tries to make an important decision about Sophia's future without consulting her.
- 2005– 43mTV-147.9 (1K)TV EpisodeDirectorDebbie AllenStarsEllen PompeoJustin ChambersChandra WilsonStephanie's relationship to Kyle becomes complicated. Meredith encourages Amelia and Owen to take a chance and choose to be a real couple. Arizona and Callie are at odds and put their friends in the middle of an uncomfortable situation.
- DirectorKevin McKiddStarsEllen PompeoJustin ChambersChandra WilsonCallie and Arizona are at odds over Sofia's future, and Stephanie questions the status of her relationship with Kyle. Meanwhile, Alex and April work on a pregnant teen that is readmitted to Grey Sloan Memorial.
- DirectorRob CornStarsEllen PompeoJustin ChambersChandra WilsonOwen and Amelia take their relationship to the next level; Alex has a clearer view of his future with Jo; Callie and Arizona struggle with custody arrangements.
- DirectorDebbie AllenStarsEllen PompeoJustin ChambersChandra WilsonMeredith and Maggie support Amelia on an important day; Jo reveals that she has been keeping secrets from Alex while Arizona and Callie deal with the impact of their custody arrangement. Meanwhile, Ben leaps into action to help a friend.
- DirectorConstantine MakrisStarsMark FeuersteinPaulo CostanzoReshma ShettyHankMed turns a new page after Boris buys Hamptons Heritage. Hank loses Charlotte and takes on the case of TV host who is hosting a party at the Newberg house. Evan and Paige discover that they may be having a baby.